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Keanu Reeves

jako: Thomas "Neo" Anderson (voice)

Tara Strong

jako: Crew Woman ("Final Flight of Osiris") / Nurse ("World Record") / Misha ("Beyond")

Tom Kenny

jako: Operator ("Final Flight of Osiris")

Carrie-Anne Moss

jako: Trinity (voice)

Kevin Michael Richardson

jako: Thadeus ("Final Flight of Osiris") / Cop ("Kid's Story") / Agent #2 ("World Record")

Tress MacNeille

jako: Housewife ("Beyond") / Kenny ("Beyond")

Olivia d'Abo

jako: Rox ("Matriculated")

Phil LaMarr

jako: Duo ("Program")

Dwight Schultz

jako: Additional Voices ("The Second Renaissance Part I & II") / Monaka ("Matriculated")

Akio Otsuka

jako: Thadeus (voice)

Kath Soucie

jako: Pudgy ("Beyond") / Masa ("Beyond") / Sara ("Beyond")

Pamela Adlon

jako: Jue ("Final Flight of the Osiris") / Manabu ("Beyond") (voice)

Victor Williams

jako: Dan ("World Record")

Debi Derryberry

jako: Kid ("The Second Renaissance Part I & II")

John DiMaggio

jako: Crew Man ("Final Flight of the Osiris") / Kaiser ("Program") (voice)

Melinda Clarke

jako: Alexa ("Matriculated") (voice)

Jill Talley

jako: Mother ("The Second Renaissance Part I & II")

Rick Gomez

jako: Pilot ("Final Flight of Osiris")

James Arnold Taylor

jako: Additional Voices ("The Second Renaissance Part I & II") / Ash ("A Detective Story") / Raul ("Matriculated")

John DeMita

jako: Teacher ("Kid's Story")

Terrence 'T.C.' Carson

jako: Clarence ("A Detective Story")

Allison Smith

jako: Reporter ("World Record")

Clayton Watson

jako: The Kid (voice)

Rodney Saulsberry

jako: Chyron ("Matriculated")

Alex Fernandez

jako: Tom ("World Record")

John Wesley

jako: Dan's Dad ("World Record")

Jack Fletcher

jako:

Hedy Burress

jako: Cis ("Program") / Yoko ("Beyond")

Bette Ford

jako: Old Woman ("Final Flight of Osiris")

Matt McKenzie

jako: Agent #1 ("World Record") / Agent ("Beyond") / Agent ("A Detective Story")

Julia Fletcher

jako: The Instructor ("The Second Renaissance Part I & II") / Narrator ("World Record")